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This is due to an impairment in dTMP synthesis which leads to cell cycle arrest in S-phase of rapidly proliferating cells, in particular hematopoietic cells

These side effects may include: Nausea Headache Fatigue Gastrointestinal discomfort While less common, more serious side effects such as irregular heartbeat and kidney problems have been reported in some cases

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subtilis , which has become a model organism among industrial riboflavin-producing strains due to its ability to secrete large amounts of protein directly into the medium in a short time (Sauer et al., 1996