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The reduced form (GSH) is biologically active and plays a critical role in neutralizing free radicals, detoxifying harmful substances, and supporting immune function
The deprotonated forms of Lys and Arg with lower pKbs are much stronger bases than the deprotonated form of His, so at physiological pH, they would always be protonated (unless their local environment lowers their pKa and pKb values)
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